Literature and Taxonomic Keys
Taxonomic literature and keys are invaluable to the benthic biologist faced with identifying fauna collected from the seabed. A guide to the published, and unpublished, taxonomic literature for North East Atlantic seas is available via the NMBAQC Bibliography of Taxonomic Literature which is updated on a regular basis. All relevant papers and keys should be listed therein. However, if you are aware of important keys that have been overlooked then please let us know. For copyright reasons we cannot make published keys available here. Many are available via open access on the internet. The NMBAQC Bibliographies are available below in (in section 1) below.
The NMBAQC scheme has itself produced or commissioned numerous taxonomic keys as outputs of its various taxonomic workshops led by recognised experts in particular taxonomic groups. Some of these have since been published as formal papers but many remain as unpublished working keys designed for everyday use in the marine laboratory. In addition, a number of unpublished in-house keys produced by the scheme contractor, or offered for use by participants of the scheme are also available. All of these current keys and guides are available (in section 2) below.
An archive of older unpublished keys has also been collated (in section 3) below. These include older NMBAQC keys, some keys by NMBAQC scheme participants, and historical keys produced at taxonomic workshops organised by the Estuarine and Coastal Sciences Association (ECSA), which was formerly known as the Estuarine and Brackish-Water Sciences Association (EBWSA). These keys go back as far as the early 1980’s and, while most have now been superceded, many are still useful. These old versions often include various hand-written annotations which have been added then or since. A browse through these keys certainly shows how the taxonomy of benthic fauna has changed over the decades, and why comparing older taxonomic records with new is potentially problematical.
1) NMBAQC Bibliographies of Taxonomic Literature
2) Current keys and guides produced by the NMBAQC scheme or in collaboration with the scheme
ANNELIDA:
Polynoidae, Polyodontidae, Sigalionidae, Pholoidae - 2011
Barnich, R. 2011. Identification of scale worms in British and Irish waters. NMBAQC 2010 taxonomic workshop, Dove Marine Laboratory. 52pp, February 2011
Pleijel, F., 2001. Family Phyllodocidae. Key to Northern European Phyllodocids. 6pp and 2 figs. IRTU/ECSA Taxonomic Workshop, Portaferry Oct. 2001, (unpublished).
Worsfold, T.M., 2007. Identification guides for the NMBAQC Scheme: 2. Goniadidae, with notes on Glyceridae (Polychaeta) from shallow seas around the British Isles. Porcupine Marine Natural History Society Newsletter, 22: 19-23.
Dnestrovskaya, N.Yu. & Jirkov, I.A. 2011. Identification key for Nephtyidae (Polychaeta) of the Eastern Atlantic and the North Polar Basin. NMBAQC 2008 taxonomic workshop, Dove Marine Laboratory. 7pp, October 2011
Jarvis, S. 2011. Hesionidae (Grube, 1850) - A Provisional Guide to the Identification of the British Species. Report to the NMBAQC 2008 taxonomic workshop participants - Dove Marine Laboratory. MIES Report, 10pp, June 2011
San Martín, G. & Worsfold, T.M. 2015. Zookeys, 488 pp.1-29.
Oug, E. 2012. Guide to identification of Lumbrineridae (Polychaeta) in north east Atlantic waters v.3.2. NMBAQC 2010 taxonomic workshop, Dove Marine Laboratory. 31pp, February 2012
Gil, J., 2016. Keys to European Paraonidae Cerruti, 1909. NMBAQC Taxonomic Workshop 2016. Millport FSC Station, 11th to 15th October 2016, 11pp., (unpublished).
Radashevsky, V.I., 2017. Identification keys and comments on the taxonomy of spionid polychaetes (Annelida: Spionidae) from the continental shelf of northern Europe. NMBAQC Ring Test no. 54, 52pp. (unpublished).
Worsfold, T.M., 2009. Progress on the identification of Cirratulidae in British and Irish waters through the NMBAQC Scheme: 1996-2009. Report to the NMBAQC Scheme. Unicomarine Report NMBAQC, cir09, July 2009, 114pp., (unpublished).
Unicomarine, 2000. A key to British Capitellidae; adapted from ECSA workshop key. Unicomarine Key, 1p. (Unpublished).
Garwood, P. R. 2007. Family Maldanidae - A guide to species in waters around the British Isles. NMBAQC 2006 taxonomic workshop, Dove Marine Laboratory.
Rowe, G. A. 2010. A Provisional Guide to the family Opheliidae (Polychaeta) from the shallow waters of the British Isles. Report to the NMBAQC 2008 taxonomic workshop participants - Dove Marine Laboratory. EMU Report, 12pp, June 2010.
Scalibregmatidae, 2006
Worsfold, T.M., 2006. Identification guides for the NMBAQC Scheme: 1. Scalibregmatidae (Polychaeta) from shallow seas around the British Isles. Porcupine Marine Natural History Society Newsletter, 20: 15-18.
Terebellomorpha, 2013,
Jirkov, I.A. and Leantovic, M.A. 2013. Identification keys for Terebellomorpha (Polychaeta) of the eastern Atlantic and the North Polar Basin. Invertebrate Zoology, 10: 217-243.
Sabellidae & Fabriciidae, 2015 - please contact Claire Taylor for this file as it is large
Giangrande, A., Licciano, M. & Wasson, B. 2015. Guide to identification of Sabellidae and Fabriciidae (Polychaeta) in north east Atlantic and Mediterranean waters. NMBAQC 2014 taxonomic workshop, Dove Marine Laboratory. 91pp, January 2015.
Worsfold, T.M., 2003. Introduction to Oligochaetes. Report to the NMBAQC 2003 Taxonomic Workshop participants - Dove Marine Laboratory. Unicomarine Report NMBAQC olig03, 22pp, November 2003
CRUSTACEA:
Guerra-García, J.M. 2014. Caprellidea. Identification guide to British caprellids v.3.3. NMBAQC 2012 Taxonomic Workshop, Dove Marine Laboratory. 17pp, September 2014.
Shalla, S.H. 2011. Cumacea - Identification guide to British cumaceans. NMBAQC 2010 Taxonomic Workshop, Dove Marine Laboratory. 46pp, July 2011
MOLLUSCA:
Brenzinger, B. 2015. Identification guide to Northern European interstitial opisthobranchs (Gastropoda: Heterobranchia). Version 2.1. NMBAQC 2014 taxonomic workshop, Dove Marine Laboratory. 23pp, August 2015.
Brenzinger, B. 2015. Identification guide to Northern European interstitial opisthobranchs (Gastropoda: Heterobranchia). Keys Appendix, Version 2.1. NMBAQC 2014 taxonomic workshop, Dove Marine Laboratory. 3pp, August 2015.
3. Archive of historical unpublished keys produced by NMBAQC, ECSA, etc.
(Order follows NMBAQC Taxonomic Bibliography 2020, but excluding keys in section above)
ANNELIDA:
Examination of Polychaete specimens,2001
Mackie, A.S.Y., 2001. Examination of polychaete specimens. 2pp. Originally from NRA Polychaete Identification Course, National Museum of Wales, Nov. 92. Re-issued at IRTU/ECSA Taxonomic Workshop, Portaferry, Oct. 2001, (unpublished).
Aphroditidae, Polynoidae, Pholoidae, Sigalionidae
Chambers, S.J., 1990. The British species of Aphroditoidea, keys to species of Polynoidae, Sigalionidae, Aphroditidae and note on Pholoe. ECSA Polychaete Workshop, Fort Popton, 1990, 15pp., (unpublished).
Chambers, S.J., 1994. Identification guide and key to Pholoe. NMBAQC Polychaete Workshop, Millport 1994, 8pp. (unpublished). Hamilton, S., 2005. North Atlantic Pholoidae (from Petersen, 1998).
Hamilton, S., 2005. North Atlantic Pholoidae (from Petersen, 1998), 2pp. (Dichotomous and tabular key – unpublished).
Loshamn, A-A. (1980). Aphroditoidae, Polynoidae, Polyodontidae & Sigalionidae. Loshamn’s key to Scandinavian and Arctic species. Translated by Fredrik Pleijel (March 1990) – from En systematik og Dryegeografisk undersokelse over skjellrygg-gruppen (Familie Aphroditidae sensu Fauvel 1923). Supplied by A.S.Y. Mackie /ECSA 1990, 8pp., (unpublished translation).
Phyllodocidae ECSA Workshop, 1990 and Phyllodocidae EBWSA Workshop, 1985
Mackie, A.S.Y., 1990. Phyllodocidae. ECSA Polychaete workshop, Fort Popton, 1990, 5pp., (unpublished). O'Connor, B.S., 1985. Phyllodocidae. EBWSA errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 6pp., (unpublished).
O'Connor, B.S., 1985. Phyllodocidae. EBWSA errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 6pp., (unpublished).
Glyceridae, Goniadidae
Glycera species of British Waters
Hunter, J., 2004. Glycera species likely to be found British waters. (unpublished).
Glyceridae and Goniadidae - EBWSA Workshop
O'Connor, B.D.S., 1985. Glyceridae and Goniadidae. EBWSA errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 7pp., (unpublished).
Keys and LIterature on Glyceridae and Goniadidae
O’Connor, B.D.S. & Worsfold, T.M., 2006. Keys to and Literature on Glyceridae and Goniadiadae. BEQUALM/NMBAQC 2006 Taxonomic Workshop, Dove Marine Lab., Nov. 2006, 19pp., (unpublished).
Unicomarine, 2000. Key to glycerid genera, key to the genus Glycera of the North-East Atlantic. 3pp. (unpublished).
Unicomarine, 2003. Key to Goniadidae
Nephtyidae
Family Nephytidae - EBWSA workshop
Garwood, P.R., 1985. Family Nephtyidae. EBWSA errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 13pp., (unpublished).
Family Nephytidae - ECSA workshop
Garwood, P.R., 1990. British Nephtyidae. ECSA polychaete workshop, Fort Popton, April 1990, 7 pp., (unpublished).
British Nephytidae - NMBAQC workshop
Garwood, P.R., 1995. British Nephtyidae. NMBAQC taxonomic workshop, Portaferry, June 1995, 22pp., (unpublished).
Family Nephytidae - Key to northern European nephytids
Pleijel, F., 2001. Family Nephtyidae. key to northern European nephtyids. 5pp and 2 figs. IRTU/ECSA Taxonomic Workshop, Portaferry Oct. 2001, (unpublished).
Key to Nephytidae - Unicomarine 2000
Unicomarine, 2000. Key to Nephtyidae. 3pp. (Unpublished)
Nereidae, Hesionidae, Pilargidae
Family Nereidae - EBSWA Workshop
Chambers, S.J., 1985. Family Nereidae. EBWSA Errant Polychaete Workshop, Heriot-Watt University, Edinburgh 1985, 13pp. (unpublished).
Key to Gyptis, Amphiduros and Podarkeopsis
Hunter, J., 2005. Key to Gyptis, Amphiduros and Podarkeopsis From Pleijel 1993. Taxonomy of European species of Amphiduros and Gyptis (Polychaeta: Hesionidae). Hunter Biological unpublished In-House key.
Mackie, A.S.Y., 1992. Key to Northern European Hesionids. Polychaete Colloquium, 1992, 4pp., (unpublished).
O'Connor, B.D.S., 1985. Hesionidae. EBWSA Errant Polychaete Workshop, Heriot-Watt University, Edinburgh, 1985, 3pp., (unpublished).
O'Connor, B.D.S., 1985. Pilargidae. EBWSA Errant Polychaete Workshop, Heriot-Watt University, Edinburgh, 1985, 2pp., (unpublished).
Unicomarine, 2003. Key to Pilargidae. 1p. (Unpublished).
Syllidae
Garwood, P.R., 1985. Family Syllidae. EBWSA Errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 25pp., (unpublished).
Garwood, P.R., 1990. Family Syllidae. ECSA polychaete workshop, Fort Popton, April 1990, 15 pp., (unpublished).
British Species of the Genus Syllis
Garwood, P.R., 1995. British species of the Genus Syllis. Identification notes. 12 pp. (unpublished).
Family Syllidae - NMBAQC Workshop
Garwood, P.R., 2006. Family Syllidae. Provisional guide to the identification of British species. NMBAQC Taxonomic Workshop, Dove Marine Lab., Nov. 2006, 34pp., (unpublished).
San Martin, G., 2013. Guide and keys for the identification of Syllidae (Polychaeta) from the British Islands (reported and expected species). NMBAQC Workshop, Dove Marine Laboratory, Cullercoats, 31pp. (Updated version published in 2015 in Zookeys).
Chambers, S.J., 1985. Family Sphaerodoridae. EBWSA errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 1pp., (unpublished).
Dorvilleidae, Lumbrineridae
Hunter, J., 2005. Some dorvilleid genera. 3pp. (Unpublished).
O'Connor, B.D.S., 1985. Eunicida. EBWSA errant polychaete workshop, Heriot-Watt University, Edinburgh, 1985, 7pp., (unpublished).
Oug, E., 2003. Characters of species of Ophryotrocha and allied forms from European waters. 6pp. NMBAQC Taxonomic Workshop, Dove Marine Lab., 2003 (Unpublished).
Lumbrineridae from NE Atlantic Waters - NMBAQC Workshop 2003
Oug, E., 2003. Lumbrineridae from north east Atlantic waters. 18pp. + 2 plates. NMBAQC Benthic Invertebrate Taxonomic Workshop, Dove Marine Laboratory, Tynemouth, UK, November 2003., (unpublished).
Paraonidae, Orbiniidae
Mackie, A.S.Y. & Hartley, J.P., 1992. The Paraonidae; key to British Paraonidae. National Rivers Association Polychaete Workshop, 10pp., (unpublished).
Key to Orbiniidae - NMBAQC Workshop
Unicomarine, 1996. Key to Orbiniidae. Key for NMBAQC Workshop, Millport, Nov. 1996, 2pp., (unpublished).
Spionidae, Magelonidae
Hall, D.J., 1995. Tentative Spio key. Unicomarine Key, 9pp. (Unpublished).
Hall, D.J., 2000. Key to Malacoceros. Unicomarine Key, 1p (Unpublished).
Hall, D.J., 2000. Key to Scolelepis (Nerinides); subgenera S. (Scolelepis) & S. (Parascolelepis). Unicomarine Key, 2pp. (Unpublished).
Hall, D.J., 2000. Tentative Spio key. Unicomarine Key, 9pp. (Unpublished).
Hall 2006 Key to Scolelepsis - updated 2000 Key
Hall, D.J., 2006. Key to Scolelepis. Updated Hall 2000 key with Scolelepis features table. Unicomarine Key, 3pp. (Unpublished).
Mackie 1990 Spionidae and Magelonidae
Mackie, A.S.Y., 1990. Spionidae and Magelonidae. ECSA polychaete workshop, Fort Popton, April 1990, 12pp., (unpublished).
Mackie A.S.Y., 2001. Key to Spionidae. 13pp. ECSA taxonomic workshop, Portaferry, Oct. 2001., (unpublished).
Mackie 2001 Key to Magelonidae
Mackie, A.S.Y., 2001. Key to Magelonidae. ECSA taxonomic workshop, Portaferry, Oct. 2001, 2pp. (unpublished).
Muir, A., 1984. British Magelonidae. EBWSA sedentary polychaete workshop, Swansea, April 1984, 2pp., (unpublished).
Radashevsky 2012 Tentative Key to Streblospio
Radashevsky, V.I., 2012. Tentative key to Streblospio. 1pp. Unpublished key for NMBAQC scheme, March 2012.
Radashevsky, V.I., 2012. Spionidae (Annelida) from shallow waters around the British Islands: an identification guide for the NMBAQC Scheme with an overview of spionid morphology and biology. Zootaxa, 3152, 1-35. *Please note this is available from the publisher (there is a more recent unpublished review of British Spionids is available - see above in current section, Radashevsky, 2017)
Radashevsky 2016 Keys to Spionidae from shallow waters around the British Isles
Radashevsky, V.I., 2016. Keys to Spionidae (Annelida) species from shallow waters around the British Islands. NMBAQC 2016 taxonomic Workshop FSC Millport, 14pp. (unpublished).
Cirratulidae
Cirratulidae Key - EBWSA workshop 1984
George, J.D., 1984. Cirratulidae Key. EBWSA sedentary polychaete workshop, Swansea, April 1984, 2pp., (unpublished).
Mackie and Chambers - Cirratulidae Key NRA Polychaete Workshop 1992
Mackie, A.S.Y. & Chambers, S.J., 1992. Cirratulidae Key. National Rivers Authority Polychaete Workshop, Nov.92, 6pp., (unpublished).
Mackie and Chambers - Cirratulidae workshop key 1994
Mackie, A.S.Y. & Chambers, S.J., 1994. Cirratulidae workshop key. Polychaete Colloquium Workshop, Newcastle, April 1994, 8pp., (unpublished).
Unicomarine - A guide to the family Cirratulidae 1996
Unicomarine Ltd., 1996. A guide to the family Cirratulidae including a key to anterior portions. Version 1.00 (RT09). Report to the NMBAQC Committee, September 1996, 79pp., (unpublished).
Unicomarine - Key to Cirratulidae Version 2.00 1998
Unicomarine Ltd., 1998. Key to Cirratulidae, (Version 2.00 , 26 Feb. 1998) 13pp., (unpublished).
Unicomarine 1998 - Notes on revised Cirratulid taxonomy Version 2 1998
Unicomarine Ltd., 1998. Notes on revised cirratulid Taxonomy for Key (v.2.00), 2pp., (unpublished).
Unicomarine - Notes to accompany species description 1998
Unicomarine Ltd., 1998. Notes to accompany the species description section of the cirratulid guide (v2.00), 4pp.,19 distribution maps, (unpublished).
Worsfold - A provisional update to the identification of UK Cirratulidae 2006
Worsfold, T.M., 2006. A provisional update to the identification of UK Cirratulidae. Report for the NMBAQC 2006 Taxonomic Workshop, Dove Marine Laboratory, 6-10 November 2006, 14pp., (unpublished).
Capitellidae, Arenicolidae, Flabelligeridae
Garwood British Capitellidae 1990
Garwood, P.R., 1990. British Capitellidae. ECSA polychaete workshop, Fort Popton, 1990, 3pp., (unpublished).
Muir, A., 1984. British lugworms. EBWSA sedentary polychaete workshop, Swansea University, April, 1984, 1pp., (unpublished).
Pleijel British Capitellidae 1982
Pleijel, F., 1982. British Capitellidae. (Unpublished).
Unicomarine Key to Flabelligeridae 2000
Unicomarine 2000 Key to Flabelligeridae. 2pp. (unpublished).
Warren Provisional Key to British Capitellids 1984
Warren, L., 1984. Provisional Key to British Capitellids. EBSWA Sedentary Polychaete Workshop, Swansea University. 5pp. (unpublished).
Maldanidae
Bamber British Maldanid Posteriors 1997
Bamber, R., 1997. British maldanid posteriors. 2pp., (unpublished tabular key).
Unicomarine Key to Maldanidae 1996
Unicomarine Ltd., 1996. Key to Maldanidae. Key for NMBAQC workshop, Millport, Nov.1996, 5pp., (unpublished).
Terebellomorpha
Garwood. Family Terebellidae: Subfamily Polycirrinae. 1994
Garwood, P.R., 1994. Family Terebellidae: Subfamily Polycirrinae. Polychaete Colloquium, Newcastle, 7pp.
Jirkov I.A., 2011. Identification keys for Terebellomorpha (Polychaeta) of the eastern Atlantic and the North Polar Basin. II. Ampharetidae. From NMBAQC 2008 Taxonomic Workshop, Dove Marine Laboratory, 6pp., (unpublished).
Leontovich, M.K. & Jirkov I.A., 2011. Identification keys for Terebellomorpha (Polychaeta) of the eastern Atlantic and the North Polar Basin. I. Pectinariidae and Terebellidae. From NMBAQC 2008 Taxonomic Workshop, Dove Marine Laboratory, 11pp., (unpublished).
Pleijel, F., 2001. Terebellomorphs. Keys to Northern European Pectinarids, Ampharetids, Terebellids, & Trichobranchids. 11 pp. IRTU/ECSA Taxonomic Workshop, Portaferry Oct.2001, (unpublished).
Sabellidae, Serpulidae (including Spirorbidae)
Knight-Jones, P., 1984. Draft keys to families Sabellidae, Serpulidae and Spirorbidae, 12, 8, & 3pp. EBWSA Sedentary Polychaete Workshop, Swansea, April 1984, (unpublished).
Mackie. Sabellidae. Key to genera. ECSA Polychaete Workshop 1990
Mackie A.S.Y., 1990. Sabellidae. Key to genera, 4pp. ECSA Polychaete Workshop, Fort Popton, 1990, (unpublished).
Mackie. Sabellidae. IRTU/ECSA Workshop 2001
Mackie A.S.Y., 2001. Key to Sabellidae. 5pp. IRTU/ECSA Taxonomic Workshop, Portaferry Oct.2001, (unpublished).
OLIOGOCHAETA
Van Haaren Oligochaetes from brackish and marine waters in The Netherlands. 2018
Van Haaren, T., 2018. Oligochaetes from brackish and marine waters in The Netherlands (Annelida:Oligochaeta), 30pp. (unpublished English translation of Van Haaren, 2016)
Unicomarine Oligochaete workshop 1994
Unicomarine, 1994. Oligochaete workshop – Preston Montford – Timetable, delegates, general notes, marine tubificid taxonomy, collection, Tubificidae reported at workshop, Tubificidae features table, V1 & V2, Preliminary key to tubificids, British species list, world species list, references, 90pp. (unpublished).
Unicomarine Tubificidae features table 1996
Unicomarine, 1996. Tubificidae features table (ver.3), 2pp. (unpublished).
Unicomarine Marine / brackish oligochaetes features table 2000
Unicomarine, 2000. Marine / brackish oligochaetes features table, 3pp. (unpublished).
ARTHROPODA
Hall Key to Coastal Species of British Mysidae 2007
Hall, D.J., 2007. Key to coastal species of British Mysidae (from Makings with Heteromysis / Heteromysoides revisions – note Heteromysis formosa is an American species). 5pp. + 2pls. (Unpublished).
Holdich & Bird British Shallow Water Isopoda – Species List and Family Key 1998
Holdich, D.M. & Bird, G., 1988. British shallow water Isopoda – species list and family key. Estuarine & Brackish-Water Sciences Association, Small Crustacean Workshop, Bangor, Wales, 1988, 4pp., (unpublished).
Holdich and Bird. British Shallow Water Tanaidacea 1988.
Holdich, D.M. & Bird, G.J., 1988. British shallow water Tanaidacea. Estuarine & Brackish-Water Sciences Association, Small Crustacean Workshop, Bangor, Wales, 1988, 19pp., (unpublished).
Moore Manual for Workshop on Amphipod Identification 1996
Moore, P.G., 1996. Manual for workshop on Amphipod Identification, Millport, Oct .96, 140pp. NMBAQC Scheme, (unpublished).
O'Reilly A Guide to Nicothoid Parasites of British Amphipods, Cumaceans and Isopods 1989
O’Reilly, M., 1989. A guide to nicothoid parasites of British amphipods, cumaceans and isopods. Estuarine & Brackish-Water Sciences Association, Small Crustacean Workshop, Bangor, Wales, 1988, (revised 1989), 14pp (unpublished).
Unicomarine Ampelisca Pictorial Key 1996
Unicomarine Ltd., 1996. Ampelisca pictorial key. 2pp. (unpublished).
MOLLUSCA
Killeen & Light. Mollusc ECSA Workshop 1994
Killeen, I., & Light, J., 1994. Mollusc Workshop - Estuarine & Coastal Sciences Association (ECSA), Heriot-Watt University, Edinburgh, April 1994. Text & figures on literature, assemblages, chitons, limpets, Trochacea, Littorinacea, Rissoacea, Hydrobia, Neogastropoda-Hinia, Conacea, Pyramidellacea, Opistobranchia, bivalves, Nuculacea-taxodonts, Mytilacea, Anomiacea, Pectinacea, Galleommatacea/Erycinacea, Lucinacea Cardiacea, Mactracea, Solenacea, Tellinacea, Veneracea, Myacea, Hiatellacea, Pandoracea, Thraciacea, 74pp., and added handwritten notes, 20pp., (unpublished). (Re-issued at IRTU/ECSA Taxonomic Workshop, Portaferry Oct. 2001).
Smith. Mollusc Taxonomic Workshop 1986
Smith, S.M., 1986. Mollusc taxonomic workshop - Estuarine and Brackish-Water Sciences Association (EBSA) – Oban April/May 1986. (Pictorial guide to British molluscs - 32 plates of collated figures copied from Tebble, Linn. Soc. Synopses etc. plus photo of 3 Abra species), (unpublished).
ECHINODERMATA
Southward. Keys for the Identification of Echinodermata of the British Isles 1972
Southward, E.C., 1972. Keys for the Identification of Echinodermata of the British Isles. Marine Biological Association - Echinoderm Survey, 24 pp., (unpublished).
Southward, E.C., & Picton, B.E., 2003. Keys for the Identification of Echinodermata of the British Isles, Marine Biological Association - Echinoderm Survey. NMBAQC Benthic Invertebrate Workshop, Dove Marine Laboratory, Cullercoats, Tynemouth, Nov.2003. 20 pp., (unpublished).
Southward & Tyler. Echinoderms EBSA Workshop 1982
Southward, E.C. & Tyler P.A., 1982. Echinoderms. Estuarine and Brackish-Water Sciences Association - (EBSA) Echinoderm Workshop, Keys and notes. University of Stirling, 7 - 10 Sep. 1982, 39pp., (unpublished).